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About | Classical Genetics | Timelines | What's New | What's Hot

About | Classical Genetics | Timelines | What's New | What's Hot


The Electronic Scholarly Publishing Project: Providing world-wide, free access to classic scientific papers and other scholarly materials, since 1993.


Recommended Videos


2016: Lectures on squid-vibrio symbiosis

Two lectures by Margaret McFall-Ngai given as part of the iBioSeminars on Microbiology

Advances in rRNA sequencing and other techniques have allowed scientists to characterize novel symbiotic partnerships. In her first lecture, Dr. Margaret McFall-Ngai provides an overview of the three main types of symbiosis: mutualism (both partners benefit), commensalism (only one partner benefit), and parasitism (one partner benefits, but the other partner is harmed). McFall-Ngai's research is currently focused on understanding the establishment and maintenance of symbiotic relationships, and the molecular effects that these relationships have on development, health, and disease.

Information Technology

2013: Big Data: Yet Another Buzzword or Actual Big Deal?

Presentation by R J Robbins at BRIITE Annual Meeting, 11-13 Dec, 2013. Salk Institute for Biological Studies, La Jolla, CA

The invention of digital information storage technology is allowing the amount of stored information on Earth to grow at an incredible rate. In 1986, 99% of stored information was in analog form; today 99.9% is digital. When n is approaching all, that is, when it is possible to access data on all relevant subjects, analyses change in both quantitative and qualitative ways. If "de-identified" data sets can be joined with other data sets, de-identification becomes impossible.


2018: Tesla Roadster + Starman in Space, Orbiting Earth

Red Car for the Red Planet

In February, 2018, Elon Musk's company, SpaceX, successfully tested its giant rocket — The Falcon Heavy — by launching a red Tesla convertible into space.

2018: Robot Dogs Walk Around, Open a Door

Nifty headless robotic dogs get stuff done.

Boston Dynamics has developed robot dogs that can walk around to investigate things. They can also, if needed, open doors. Cool. They will be going on sale soon.

ESP Quick Facts

ESP Origins

In the early 1990's, Robert Robbins was a faculty member at Johns Hopkins, where he directed the informatics core of GDB — the human gene-mapping database of the international human genome project. To share papers with colleagues around the world, he set up a small paper-sharing section on his personal web page. This small project evolved into The Electronic Scholarly Publishing Project.

ESP Support

In 1995, Robbins became the VP/IT of the Fred Hutchinson Cancer Research Center in Seattle, WA. Soon after arriving in Seattle, Robbins secured funding, through the ELSI component of the US Human Genome Project, to create the original ESP.ORG web site, with the formal goal of providing free, world-wide access to the literature of classical genetics.

ESP Rationale

Although the methods of molecular biology can seem almost magical to the uninitiated, the original techniques of classical genetics are readily appreciated by one and all: cross individuals that differ in some inherited trait, collect all of the progeny, score their attributes, and propose mechanisms to explain the patterns of inheritance observed.

ESP Goal

In reading the early works of classical genetics, one is drawn, almost inexorably, into ever more complex models, until molecular explanations begin to seem both necessary and natural. At that point, the tools for understanding genome research are at hand. Assisting readers reach this point was the original goal of The Electronic Scholarly Publishing Project.

ESP Usage

Usage of the site grew rapidly and has remained high. Faculty began to use the site for their assigned readings. Other on-line publishers, ranging from The New York Times to Nature referenced ESP materials in their own publications. Nobel laureates (e.g., Joshua Lederberg) regularly used the site and even wrote to suggest changes and improvements.

ESP Content

When the site began, no journals were making their early content available in digital format. As a result, ESP was obliged to digitize classic literature before it could be made available. For many important papers — such as Mendel's original paper or the first genetic map — ESP had to produce entirely new typeset versions of the works, if they were to be available in a high-quality format.

ESP Help

Early support from the DOE component of the Human Genome Project was critically important for getting the ESP project on a firm foundation. Since that funding ended (nearly 20 years ago), the project has been operated as a purely volunteer effort. Anyone wishing to assist in these efforts should send an email to Robbins.

ESP Plans

With the development of methods for adding typeset side notes to PDF files, the ESP project now plans to add annotated versions of some classical papers to its holdings. We also plan to add new reference and pedagogical material. We have already started providing regularly updated, comprehensive bibliographies to the ESP.ORG site.

Electronic Scholarly Publishing
961 Red Tail Lane
Bellingham, WA 98226

E-mail: RJR8222 @

Papers in Classical Genetics

The ESP began as an effort to share a handful of key papers from the early days of classical genetics. Now the collection has grown to include hundreds of papers, in full-text format.

Digital Books

Along with papers on classical genetics, ESP offers a collection of full-text digital books, including many works by Darwin and even a collection of poetry — Chicago Poems by Carl Sandburg.


ESP now offers a large collection of user-selected side-by-side timelines (e.g., all science vs. all other categories, or arts and culture vs. world history), designed to provide a comparative context for appreciating world events.


Biographical information about many key scientists (e.g., Walter Sutton).

Selected Bibliographies

Bibliographies on several topics of potential interest to the ESP community are automatically maintained and generated on the ESP site.

ESP Picks from Around the Web (updated 07 JUL 2018 )